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Mail routing btween Source and Target

Good Morning,

I have a question regarding mail routing between Source and Target domains.

I'd like to know how the mail flow works after  additional addresses (source.local and target.local) are set up.

Does Qmm proxy mail between domains or the flow it's done between the Exchange environments with no qmmex working in the middle of the mail routing?

Basically is it SOURCE <--> TARGET or SOURCE <--QMMEX--> TARGET

Thank you

Slav